Many of these are fantastic, I could swear some of them used actual animation. For example http://www.psy.ritsumei.ac.jp/~akitaoka/kagero3e.html
What causes the appearance of movement?
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I suspect it's your brain trying to reconcile misleading shadow/highlight information.
I thought it was misrepresentating the shadows and highlights as the "negative" afterglow you get after something has moved on the retina.